Clad in desert pink and the size of a mere child, the infamous "Pink Devil" mercilessly hunts down other players in the firearm-centered world of the virtual reality game Gun Gale Online. But in real life, this feared player killer is not quite who anyone would expect. A shy university student in Tokyo, Karen Kohiruimaki stands in stark contrast to her in-game avatar—in fact, she happens to stand above everyone else too, much to her dismay. Towering above all the people around her, Karen's insecurities over her height reach the point where she turns to the virtual world for an escape. Starting game after game in hopes of manifesting as a cute, short character, she finally obtains her ideal self in the world of Gun Gale Online. Overjoyed by her new persona, she pours her time into the game as LLENN, garnering her reputation as the legendary player killer. However, when one of LLENN's targets gets the best of her, she ends up meeting Pitohui, a skilled yet eccentric woman. Quickly becoming friends with Karen, Pitohui insists that LLENN participates in Squad Jam, a battle royale that pits teams against one another, fighting until only one remains. Thrust into the heated competition, LLENN must fight with all her wit and will if she hopes to shoot her way to the top.

The social and political reforms implemented by Reinhard are finding growing consensus among the citizens of the Galactic Empire. Meanwhile, the Free Planets Alliance is experiencing a declining phase, its national power weakened significantly due to widespread government corruption, despite military success has been achieved on the battlefields. As the balance of power between the two super states dominating the Galaxy is clearly undergoing a major shift, Adrian Rubinsky, the Landesherr of the autonomous and nominally neutral dominion of Fezzan, sees an opportunity to take advantage from the situation.

Taking place one year before the Zentraedi arrive on Earth, Macross Zero chronicles the final days of the war between the U.N. Spacy and anti-U.N. factions. After being shot down by the anti-U.N.'s newest fighter plane, ace pilot Shin Kudo finds himself on the remote island of Mayan, where technology is almost non-existent. While Shin stays on the island to heal his wounds, the tranqulity of the island is shattered by a battle that involves the UN's newest fighter - the VF-0.

After warping light-years away from Earth, the spacecraft Macross carries a city of civilians and soldiers back home. They continuously fight off the threat of the Zentradi and Meltrandi, giant alien races with whom humanity wages war. During a battle, the Macross transforms into a mecha operated by fighter pilot Hikaru Ichijou. Thrust into a number of hardships, Hikaru grows close to the city's idol Minmay Lynn. As their relationship develops, however, it is viewed unfavorably by others due to Minmay's status and popularity. With war raging on and the state of Earth unknown, Hikaru and his fellow soldiers fight on against their mysterious enemies. Throughout the bloodshed, Hikaru and Minmay attempt to keep hold of their feelings for one another. Minmay pours her heart into her music, which may just turn out to be the key to finding peace.