What do an alien, former ninja, weak telepath, and boy pretending to be a girl have in common? They are all classmates at Kashihara Girls' Academy, and they hide their true identities from the world. Rikka, Chiyo, Sekine, and Tsubasa seem like the best of friends, but they each harbor a monumental secret that they cannot reveal, no matter what. In order to continue masquerading as normal middle school girls, each of them must invent creative ways to conceal their true selves. But that is easier said than done, as the four students end up entangled in numerous shenanigans along the way.

Kind-hearted Himuro-kun is unfortunate: when in deep concentration, stressed, or flustered, he involuntarily creates blizzard conditions for himself and anyone in his vicinity. During one such incident on the first day of his new job, Himuro-kun encounters the beautiful Fuyutsuki-san, who helps him break out of his nerve-induced ice. As it turns out, Fuyutsuki-san is his new coworker. At the office, the cool-headed Fuyutsuki-san offers simple and rational solutions to Himuro-kun's icy dilemmas—everything from helping him garden without freezing his plants to ensuring he does not melt during their tropical work retreat. Every time Fuyutsuki-san does something kind for him, the tempest of emotions he experiences inside embarrassingly manifests on the outside. As a result of the frequent snowstorms, Himuro-kun's feelings for Fuyutsuki-san are impossible to hide. Even though Fuyutsuki-san is unfamiliar with love, Himuro-kun remains determined to repay her kindness and warm her heart in any way he can.

Mitsuha Yamano could not have had it worse. After failing to enter her university of choice and losing her entire family in an accident, she finds herself plummeting off a cliff due to an especially fragile guardrail. Through what seems to be the only bit of the fortune left in her life, Mitsuha's firm unwillingness to die allows her to be transported to another world, where she ends up surviving. In this foreign land, Mitsuha wanders around for days before collapsing from exhaustion and being rescued by a little girl named Colette. Not long after, she finds herself in yet another life-threatening situation, and her indomitable will shines once more, granting her an audience with a divine cosmic being. The entity reveals to Mitsuha that she has acquired not only superhuman regeneration but also the ability to transport between worlds at will. Armed with this new information, Mitsuha brainstorms ways to utilize her newfound power to its full potential. Realizing that she can take goods from Earth and resell them to the other world at a higher price, she resolves to save up eighty thousand gold coins to be able to retire comfortably.

Despite the prefectural tournament's breathtaking conclusion, the Kazemai High School Kyudo Club has no time to celebrate. With the national competition right around the corner, the club is determined to prove that their initial victory was not a mere fluke. With his passion for the sport reignited, Minato Narumiya earnestly prepares for the upcoming competition with his teammates, who are equally motivated to win. As the much-anticipated tournament begins, the members of Kazemai's kyudo team fight to stay on target with their ambitious goal. But when old acquaintances reappear as difficult opponents, the boys find that winning the nationals will require more dedication and effort than ever before.